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we might have broken his trust by handing him over to the vet
I would say don't fret over that, dogs don't really bear grudges in that way.
play biting their hands.
It's more likely he wasn't taught bite inhibition. Puppies use their mouths in play with their siblings so when they come to live with their humans they normally are taught that this is no longer ok. Try keeping a toy handy to put in his mouth so he can't mouth the hand, or as soon as teeth get skin ask the person to firmly turn and walk out of the room. If you are out with him, immediately walk him away. The objective is for him to learn that teeth on skin = end of play.